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Livermore
Livermore
ZHEN

ProShare Advisors LLC

SEC CIK
0001357955
Latest filing
Aug 11, 2026

Holdings in Q1 2022

Holdings as of Mar 31, 2022, filed May 10, 2022. Long positions only; changes compare share counts with Q4 2021.

Institution overview
Positions
2,196
−142 vs. Q4 2021
Portfolio value
$46.6B
−$4.41B (−8.6%) vs. Q4 2021
Filed
May 10, 2022
SEC
Holdings of ProShare Advisors LLC in Q1 2022
SecurityClassSharesValue% of portfolioShare changeActionHistory
Ozon Hldgs PLC69269L104SPONSORED ADS63,825$00.0%+14,435+29.2%Added–
AVTXAvalo Therapeutics, Inc.COM11,679$8.00K<0.1%−1,931−14.2%ReducedHistory
LVOLiveOne, Inc.COMMON STOCK11,422$9.00K<0.1%−1,466−11.4%ReducedHistory
HOFVHall of Fame Resort & Entertainment CoCOM10,444$12.0K<0.1%−1,730−14.2%ReducedHistory
MREOMereo BioPharma Group plcSPON ADS12,056$14.0K<0.1%−3,590−22.9%ReducedHistory
AWHLAspira Women's Health Inc.COM13,629$14.0K<0.1%−2,258−14.2%ReducedHistory
UAVSAgEagle Aerial Systems Inc.COM13,081$15.0K<0.1%−1,741−11.7%ReducedHistory
LITSLite Strategy, Inc.COM NEW23,975$15.0K<0.1%+287+1.2%AddedHistory
NBEVNewAge, Inc.COM25,222$15.0K<0.1%−4,169−14.2%ReducedHistory
SEELSeelos Therapeutics, Inc.COM18,419$15.0K<0.1%−3,043−14.2%ReducedHistory
TECXTectonic Therapeutic, Inc.COM11,620$16.0K<0.1%−2,803−19.4%ReducedHistory
AMPEAmpio Pharmaceuticals, Inc.COM36,533$17.0K<0.1%−6,032−14.2%ReducedHistory
IBIOiBio, Inc.COM NEW40,522$17.0K<0.1%−6,687−14.2%ReducedHistory
TRVNTrevena IncCOM30,661$17.0K<0.1%−5,064−14.2%ReducedHistory
KLTRKaltura IncCOM10,307$18.0K<0.1%+10,307NewHistory
LOTZCarLotz, Inc.COM CL A13,420$18.0K<0.1%−2,215−14.2%ReducedHistory
FBIOFortress Biotech, Inc.COM13,758$18.0K<0.1%−2,504−15.4%ReducedHistory
PAVMPAVmed Inc.COM13,626$18.0K<0.1%−2,256−14.2%ReducedHistory
QTNTQuotient LtdSHS14,924$18.0K<0.1%−2,218−12.9%ReducedHistory
ARDXArdelyx, Inc.COM18,117$19.0K<0.1%−2,991−14.2%ReducedHistory
VHCVirnetX Holding CorpCOM11,890$19.0K<0.1%−1,974−14.2%ReducedHistory
MBIIPro Farm Group, Inc.COM18,983$20.0K<0.1%−3,136−14.2%ReducedHistory
NGNENeurogene Inc.COM10,990$20.0K<0.1%−2,662−19.5%ReducedHistory
SYRSSyros Pharmaceuticals, Inc.COM17,205$21.0K<0.1%−4,098−19.2%ReducedHistory
ATNXAthenex, Inc.COM27,583$22.0K<0.1%−6,705−19.6%ReducedHistory
Ernexa Therapeutics Inc.114082100COM10,946$22.0K<0.1%−2,867−20.8%Reduced–
AQBAquabounty Technologies IncCOM NEW12,275$23.0K<0.1%+810+7.1%AddedHistory
MBIOMustang Bio, Inc.COM23,539$23.0K<0.1%−5,368−18.6%ReducedHistory
ATHXAthersys, IncCOM39,922$24.0K<0.1%−4,817−10.8%ReducedHistory
DSDrive Shack Inc.COM15,489$24.0K<0.1%−2,561−14.2%ReducedHistory
SIENProject Sage Oldco, Inc.COM10,885$24.0K<0.1%−1,799−14.2%ReducedHistory
NMTR9 Meters Biopharma, Inc.COM42,452$26.0K<0.1%−7,010−14.2%ReducedHistory
QMCOQuantum CorpCOM NEW11,111$26.0K<0.1%−1,660−13.0%ReducedHistory
SFTGQShift Technologies, Inc.CL A11,268$26.0K<0.1%−2,296−16.9%ReducedHistory
SLDBSolid Biosciences Inc.COM22,591$27.0K<0.1%−5,969−20.9%ReducedHistory
WVEWave Life Sciences, Inc.SHS14,386$28.0K<0.1%−3,568−19.9%ReducedHistory
ASXCAsensus Surgical, Inc.COM43,870$28.0K<0.1%−7,248−14.2%ReducedHistory
ATOSAtossa Therapeutics, Inc.COM21,936$28.0K<0.1%−3,624−14.2%ReducedHistory
RMORomeo Power, Inc.COM18,342$28.0K<0.1%−9,913−35.1%ReducedHistory
PBYIPuma Biotechnology, Inc.COM10,284$29.0K<0.1%−2,523−19.7%ReducedHistory
ADAPYAdaptimmune Therapeutics PLCSPONDS ADR14,203$29.0K<0.1%−5,112−26.5%ReducedHistory
SYRESpyre Therapeutics, Inc.COM12,680$29.0K<0.1%−3,091−19.6%ReducedHistory
DRRXDurect CorpCOM42,415$29.0K<0.1%−7,006−14.2%ReducedHistory
INFIInfinity Pharmaceuticals, Inc.COM25,623$29.0K<0.1%−6,038−19.1%ReducedHistory
LTRPALiberty TripAdvisor Holdings, Inc.COM SER A13,659$29.0K<0.1%−2,258−14.2%ReducedHistory
MILEMetromile, LLCCOM21,381$29.0K<0.1%+435+2.1%AddedHistory
NCMINational CineMedia, Inc.COM11,269$29.0K<0.1%−1,864−14.2%ReducedHistory
IMDXInsight Molecular Diagnostics Inc.COM20,825$31.0K<0.1%−5,311−20.3%ReducedHistory
Maiden Holdings, Ltd.G5753U112SHS13,032$32.0K<0.1%−2,157−14.2%Reduced–
SURFSurface Oncology, Inc.COM11,399$33.0K<0.1%−2,817−19.8%ReducedHistory
TNXPTonix Pharmaceuticals Holding Corp.COM145,212$33.0K<0.1%−13,548−8.5%ReducedHistory
ADVMAdverum Biotechnologies, Inc.COM26,444$34.0K<0.1%−6,358−19.4%ReducedHistory
DNTHDianthus Therapeutics, Inc.COM11,691$34.0K<0.1%−3,121−21.1%ReducedHistory
TLMDSOC Telemed, Inc.COM CL A11,385$34.0K<0.1%−1,884−14.2%ReducedHistory
CARMCarisma Therapeutics Inc.COM58,386$34.0K<0.1%−13,682−19.0%ReducedHistory
VIEWView, Inc.COM CL A18,396$34.0K<0.1%−12,043−39.6%ReducedHistory
AKBAAkebia Therapeutics, Inc.COM50,807$36.0K<0.1%−11,934−19.0%ReducedHistory
LCTXLineage Cell Therapeutics, Inc.COM23,513$36.0K<0.1%−3,884−14.2%ReducedHistory
XERSXeris Biopharma Holdings, Inc.COM13,983$36.0K<0.1%−3,578−20.4%ReducedHistory
RNACCartesian Therapeutics, Inc.COM29,330$36.0K<0.1%−7,209−19.7%ReducedHistory
ADMAAdma Biologics, Inc.COM20,265$37.0K<0.1%−7,311−26.5%ReducedHistory
SMMTSummit Therapeutics Inc.COM15,054$37.0K<0.1%−4,464−22.9%ReducedHistory
EVLOEvelo Biosciences, Inc.COM11,224$37.0K<0.1%−2,938−20.7%ReducedHistory
VLDRVelodyne Lidar, Inc.COM14,351$37.0K<0.1%−2,160−13.1%ReducedHistory
OTLKOutlook Therapeutics, Inc.COM NEW21,356$38.0K<0.1%+1,895+9.7%AddedHistory
PASGPassage BIO, Inc.COM12,554$38.0K<0.1%−3,170−20.2%ReducedHistory
VKTXViking Therapeutics, Inc.COM12,825$39.0K<0.1%−2,122−14.2%ReducedHistory
SCORComscore, Inc.COM13,129$39.0K<0.1%−2,167−14.2%ReducedHistory
TCRTAlaunos Therapeutics, Inc.COM61,668$41.0K<0.1%−14,568−19.1%ReducedHistory
TKTeekay Corp LtdCOM13,020$41.0K<0.1%−2,153−14.2%ReducedHistory
CTXRCitius Pharmaceuticals, Inc.COM NEW23,231$42.0K<0.1%−1,574−6.3%ReducedHistory
QTTBQ32 Bio Inc.COM13,776$42.0K<0.1%−3,438−20.0%ReducedHistory
RBBNRibbon Communications Inc.COM13,884$43.0K<0.1%−2,142−13.4%ReducedHistory
PRTKParatek Pharmaceuticals, Inc.COM14,375$43.0K<0.1%−3,383−19.1%ReducedHistory
MTEMMolecular Templates, Inc.COM12,760$44.0K<0.1%−3,249−20.3%ReducedHistory
SPRUSpruce Power Holding CorpCOM CL A22,479$44.0K<0.1%+22,479NewHistory
ZYZymergen Inc.COM15,032$44.0K<0.1%−2,432−13.9%ReducedHistory
TXMDTherapeuticsMD, Inc.COM117,101$45.0K<0.1%−27,912−19.2%ReducedHistory
VTGNVistagen Therapeutics, Inc.COM NEW36,378$46.0K<0.1%−6,013−14.2%ReducedHistory
DTILPrecision Biosciences IncCOM15,764$48.0K<0.1%−3,834−19.6%ReducedHistory
EPZMEpizyme, Inc.COM43,486$50.0K<0.1%+8,868+25.6%AddedHistory
CTMXCytomX Therapeutics, Inc.COM18,904$51.0K<0.1%−4,418−18.9%ReducedHistory
IVVDInvivyd, Inc.COM11,037$51.0K<0.1%+11,037NewHistory
LABStandard Biotools Inc.COM14,283$51.0K<0.1%−2,363−14.2%ReducedHistory
VIRXViracta Therapeutics, Inc.COM10,656$51.0K<0.1%−2,518−19.1%ReducedHistory
PSTXPoseida Therapeutics, Inc.COM11,861$53.0K<0.1%−3,223−21.4%ReducedHistory
TOONKartoon Studios, Inc.COM52,922$54.0K<0.1%−8,737−14.2%ReducedHistory
NAUTNautilus Biotechnology, Inc.COM12,877$56.0K<0.1%−4,608−26.4%ReducedHistory
SBSafe Bulkers, Inc.COM11,880$56.0K<0.1%−1,968−14.2%ReducedHistory
URGUr-Energy IncCOM34,763$56.0K<0.1%−4,772−12.1%ReducedHistory
ARAYAccuray IncCOM17,246$57.0K<0.1%−2,445−12.4%ReducedHistory
QSIQuantum-Si IncCOM CL A12,158$57.0K<0.1%−4,385−26.5%ReducedHistory
VERUVeru Inc.COM12,076$58.0K<0.1%−1,996−14.2%ReducedHistory
VBIVVBI Vaccines Inc/BCCOM NEW35,338$59.0K<0.1%−5,835−14.2%ReducedHistory
LXRXLexicon Pharmaceuticals, Inc.COM NEW28,535$60.0K<0.1%−7,722−21.3%ReducedHistory
LENZLENZ Therapeutics, Inc.COM11,546$60.0K<0.1%−184−1.6%ReducedHistory
SPPISpectrum Pharmaceuticals IncCOM47,538$61.0K<0.1%−16,307−25.5%ReducedHistory
NPKINPK International Inc.COM SHS16,933$62.0K<0.1%−2,802−14.2%ReducedHistory
CRISCuris IncCOM NEW25,720$62.0K<0.1%−6,098−19.2%ReducedHistory
GRTSGritstone bio, Inc.COM15,041$62.0K<0.1%−3,871−20.5%ReducedHistory

Sold out since Q4 2021 (171)

Positions held at the end of Q4 2021 that are gone from this filing.

The largest 100 of 171 by value last quarter.

Positions of ProShare Advisors LLC sold out since Q4 2021
SecurityClassShares in Q4 2021Value in Q4 2021% of portfolio in Q4 2021History
XLNXXilinx IncCOM517,742$109.8M0.2%History
PTONPeloton Interactive, Inc.CL A COM499,520$17.9M<0.1%History
INFOIHS Markit Ltd.SHS78,709$10.5M<0.1%History
BMTCBryn Mawr Bank CorpCOM217,151$9.77M<0.1%History
CONECyrusOne Holdco LLCCOM32,455$2.91M<0.1%History
ARNAArena Pharmaceuticals IncCOM NEW22,148$2.06M<0.1%History
CITCit Group IncCOM NEW30,668$1.57M<0.1%History
NUANNuance Communications, Inc.COM20,551$1.14M<0.1%History
HFCHollyFrontier CorpCOM33,312$1.09M<0.1%History
Athene Holding Ltd.G0684D107CL A11,272$939.0K<0.1%–
FMBIFirst Midwest Bancorp IncCOM28,472$583.0K<0.1%History
MGLNMagellan Health IncCOM NEW5,923$563.0K<0.1%History
VCRAVocera Communications, Inc.COM7,538$488.0K<0.1%History
GWBGreat Western Bancorp, Inc.COM13,829$470.0K<0.1%History
STLSterling BancorpCOM17,420$449.0K<0.1%History
Monmouth Real Estate Investment Corp609720107CL A20,817$438.0K<0.1%–
CFLTConfluent, Inc.CLASS A COM4,987$380.0K<0.1%History
GDYNGrid Dynamics Holdings, Inc.CL A9,932$377.0K<0.1%History
CMRCCommerce.com, Inc.COM SER 110,570$373.0K<0.1%History
KRAKraton CorpCOM7,875$366.0K<0.1%History
BANDBandwidth Inc.COM CL A5,043$361.0K<0.1%History
AAMIAcadian Asset Management Inc.COM13,794$353.0K<0.1%History
TXG10x Genomics, Inc.CL A COM2,300$343.0K<0.1%History
PEGAPegasystems IncCOM2,939$329.0K<0.1%History
ZGNXZogenix, Inc.COM NEW20,135$327.0K<0.1%History
RAPTRAPT Therapeutics, Inc.COM8,816$324.0K<0.1%History
FSLYFastly, Inc.CL A8,809$312.0K<0.1%History
HZOMarinemax IncCOM5,291$312.0K<0.1%History
NBPNovaBridge BiosciencesSPONSORED ADS6,553$311.0K<0.1%History
PACKRanpak Holdings Corp.COM CL A8,219$309.0K<0.1%History
ARCTArcturus Therapeutics Holdings Inc.COM8,322$308.0K<0.1%History
VREXVarex Imaging CorpCOM9,610$304.0K<0.1%History
PARPar Technology CorpCOM5,517$291.0K<0.1%History
ACCDAccolade, Inc.COM10,996$289.0K<0.1%History
QTRXQuanterix CorpCOM6,743$286.0K<0.1%History
HCMHUTCHMED (China) LtdSPONSORED ADS8,122$285.0K<0.1%History
ANABAnaptysbio, IncCOM8,085$282.0K<0.1%History
PLCEChildrens Place, Inc.COM3,514$279.0K<0.1%History
SRRKScholar Rock Holding CorpCOM11,116$277.0K<0.1%History
BALYBally's CorpCOM7,136$272.0K<0.1%History
SMPStandard Motor Products, Inc.Stock5,171$270.0K<0.1%History
AMWDAmerican Woodmark CorpCOM4,141$270.0K<0.1%History
OPRXOptimizeRx CorpCOM NEW4,312$269.0K<0.1%History
HIBBHibbett IncCOM3,712$267.0K<0.1%History
WRLDWorld Acceptance CorpCOM1,077$264.0K<0.1%History
GMABGenmab A/SSPONSORED ADS6,608$261.0K<0.1%History
ANGOAngiodynamics IncCOM9,403$259.0K<0.1%History
ZUMZZumiez IncCOM5,333$256.0K<0.1%History
UIUbiquiti Inc.COM819$251.0K<0.1%History
CCRNCross Country Healthcare IncCOM8,954$249.0K<0.1%History
BRBRBellring Brands, Inc.COM CL A8,699$248.0K<0.1%History
WOWWideOpenWest, Inc.COM11,451$246.0K<0.1%History
HYFMHydrofarm Holdings Group, Inc.COM8,510$241.0K<0.1%History
XMQualtrics International Inc.COM CL A6,806$241.0K<0.1%History
ASPNAspen Aerogels IncCOM4,819$240.0K<0.1%History
BHEBenchmark Electronics IncCOM8,794$239.0K<0.1%History
PDFSPDF Solutions IncCOM7,493$239.0K<0.1%History
EGRXEagle Pharmaceuticals, Inc.COM4,674$238.0K<0.1%History
GESGuess IncCOM10,019$238.0K<0.1%History
HURNHuron Consulting Group Inc.Stock4,739$236.0K<0.1%History
AGYSAgilysys IncCOM5,254$234.0K<0.1%History
MATVMativ Holdings, Inc.COM7,830$234.0K<0.1%History
FDMT4D Molecular Therapeutics, Inc.COM10,626$233.0K<0.1%History
MODNModel N, Inc.COM7,756$233.0K<0.1%History
DMCDel Monte CorpORD8,371$232.0K<0.1%History
Sapiens Intl Corp N VG7T16G103SHS6,762$232.0K<0.1%–
LASRNlight, Inc.COM9,532$229.0K<0.1%History
MCBMetropolitan Bank Holding Corp.COM2,139$228.0K<0.1%History
JYNTJOINT CorpCOM3,472$228.0K<0.1%History
XENEXenon Pharmaceuticals Inc.COM7,266$227.0K<0.1%History
PETQPetIQ, Inc.COM CL A9,984$227.0K<0.1%History
BATRKAtlanta Braves Holdings, Inc.COM C BRAVES GRP7,989$225.0K<0.1%History
STOKStoke Therapeutics, Inc.COM9,346$224.0K<0.1%History
NCNOnCino, Inc.COM4,091$224.0K<0.1%History
AKROAkero Therapeutics, Inc.COM10,529$223.0K<0.1%History
GRCGorman Rupp CoCOM4,965$222.0K<0.1%History
ECHOEchoStar CORPCL A8,384$221.0K<0.1%History
SCSCScansource, Inc.COM6,298$220.0K<0.1%History
PLYMPlymouth Industrial REIT, Inc.COM6,835$219.0K<0.1%History
PRSUPursuit Attractions & Hospitality, Inc.COM5,084$217.0K<0.1%History
NTGRNetgear, Inc.COM7,410$217.0K<0.1%History
CSVCarriage Services IncCOM3,357$217.0K<0.1%History
BLNKBlink Charging Co.COM7,981$212.0K<0.1%History
HFWAHeritage Financial CorpCOM8,681$212.0K<0.1%History
GABCGerman American Bancorp, Inc.Stock5,396$211.0K<0.1%History
HSIIHeidrick & Struggles International IncCOM4,839$211.0K<0.1%History
CLFDClearfield, Inc.COM2,490$210.0K<0.1%History
CALCaleres IncCOM9,251$209.0K<0.1%History
GOODGladstone Commercial CorpCOM8,075$209.0K<0.1%History
KNTEKinnate Biopharma Inc.COM11,778$209.0K<0.1%History
OBKOrigin Bancorp, Inc.COM4,836$208.0K<0.1%History
COWNCowen Inc.CL A NEW5,765$208.0K<0.1%History
NXQuanex Building Products CORPCOM8,368$207.0K<0.1%History
DGIIDigi International IncCOM8,446$207.0K<0.1%History
NTRANatera, Inc.COM2,209$206.0K<0.1%History
ALXOAlx Oncology Holdings IncCOM9,579$205.0K<0.1%History
AMTBAmerant Bancorp Inc.CL A5,902$205.0K<0.1%History
LBRDKLiberty Broadband CorpCOM SER C1,277$205.0K<0.1%History
GRBKGreen Brick Partners, Inc.COM6,719$204.0K<0.1%History
ROKURoku, IncCOM CL A895$204.0K<0.1%History